Diagnosis of genetic diseases with immune or neurological dysfunction
Submitting Institution
University of SussexUnit of Assessment
Biological SciencesSummary Impact Type
TechnologicalResearch Subject Area(s)
Biological Sciences: Biochemistry and Cell Biology, Genetics
Summary of the impact
The Caldecott/Jeggo/O'Driscoll laboratories have identified human genetic
diseases that are
caused by defects in genes involved in DNA strand-break repair. Many of
these diseases are
associated with neurological pathologies such as cerebellar ataxia
(resulting in poor balance,
movement control, and patients often being wheelchair bound), microcephaly
(smaller-than-normal
head circumference), and developmental delay. The
Caldecott/Jeggo/O'Driscoll laboratories have
engaged in identifying/diagnosing patients with such diseases as a service
to clinicians/clinical
geneticists in the UK National Health Service (NHS) and worldwide. Since
2008, these laboratories
have identified the underlying genetic defect in more than 150 patients
with a range of hereditary
DNA damage-related disorders. In particular, these laboratories have
diagnosed patients with
genetic defects in the DNA damage response genes Lig4, NHEJ1-XLF,
DCLRE1C-Artemis,
PRKDC-DNA-PKcs, PCNT, ORC1, ATRIP, ATR, and TDP2.
These diagnoses benefit both the
clinical geneticist and the patient; identifying not only the cause of the
patient's disease but also
enabling better disease management. For example, if not first diagnosed,
standard
chemotherapeutic regimes can be fatal in cancer patients who harbour
homozygous TDP2
mutations, and standard conditioning regimes used during bone-marrow
transplantation can be
fatal in LIG4 Syndrome patients. These diagnoses can therefore translate
into increased patient
survival.
Underpinning research
Neurological disease is crippling and often life-threatening, and can
result from a variety of
hereditary genetic defects. Some of these defects reflect an inability of
the affected individuals to
process and/or repair DNA damage effectively. As a result of their work
into fundamental
mechanisms of DNA damage-response and repair, the
Caldecott/Jeggo/O'Driscoll laboratories
have identified a number of human genes that, if mutated, result in
neurodegeneration and/or
neurodevelopmental defects.
For example, the Caldecott laboratory has identified TDP1, APTX,
APLF and TDP2 as novel
human genes involved in the repair of DNA single- and double-strand
breaks, and has established
that mutations in three of these cause neurodegenerative disease [see
Section 3, R1-R4].
Intriguingly, three of these four genes are required for a single stage of
DNA repair, in which the
ends of DNA breaks are processed and chemically `tidied' in readiness for
their rejoining.
Similarly, the O'Driscoll and Jeggo laboratories have identified genetic
defects that cause
microcephaly (in which affected individuals possess a smaller-than-normal
head circumference)
and dwarfism, including Seckel Syndrome (due to mutations in the genes ATR
and ATRIP),
Microcephalic Osteodysplasic Primordial Dwarfism type-II (due to mutations
in the gene PCNT)
and Meier-Gorlin Syndrome (due to mutations in ORC1) [R5-R7].
These genes are normally
involved in regulating cell duplication and division, and so result in
growth defects if mutated.
In addition, the Jeggo laboratory has identified the genetic basis of
disorders associated with
combined immunodeficiency and with elevated sensitivity to ionising
radiation (RS-SCID), such as
LIG4 Syndrome (due to mutations in Lig4) [R8].
These genes are central components of the biochemical pathway by which
chromosomal double-
strand breaks are repaired; a process that is critical to the development
of the immune system and
to resistance to X-rays and some types of chemotherapy. As a direct result
of these fundamental
discoveries in the field of DNA damage-response and repair, these
laboratories are now frequently
approached by clinical geneticists in the UK and abroad for advice and for
diagnostic input, with
the aim of identifying the underlying genetic cause of hereditary disease
in which specific DNA
damage-response mechanisms are defective.
Key researchers
- Professor Keith Caldecott, Deputy Director of the Genome Damage and
Stability Centre,
University of Sussex, 2002-present.
- Dr Mark O'Driscoll, Principal Investigator, Genome Damage and
Stability Centre, University of
Sussex, 1999-present.
- Professor Penelope Jeggo, Principal Investigator, Genome Damage and
Stability Centre,
University of Sussex, 1988-present.
References to the research
R1 Ahel, I., Rass, U., El-Khamisy, S.F., Katyal, S., Clements,
P.M., McKinnon, P.J., Caldecott,
K.W. and West, S.C. (2006) `The neurodegenerative disease protein
aprataxin resolves
abortive DNA ligation intermediates', Nature, 443(7112): 713-16.
R2 Cortes Ledesma, F., El-Khamisy, S.F., Zuma, M.C., Osborn, K.
and Caldecott, K.W. (2009)
`A human 5'-tyrosyl DNA phosphodiesterase that repairs
topoisomerase-mediated DNA
damage', Nature, 461(7264): 674-8.
R3 El-Khamisy, S.F., Saifi Gulam, M., Weinfeld, M., Helleday, T.,
Lupski., J.R. and Caldecott,
K.W. (2005) `Defective DNA single-strand break repair in spinocerebellar
ataxia with axonal
neuropathy-1', Nature, 434(7029): 108-13.
R4 O'Driscoll, M., Ruiz-Perez, V.L., Woods, C.G., Jeggo, P.A. and
Goodship, J.A. (2003) `A
splicing mutation affecting expression of ataxia-telangiectasia and
Rad3-related protein
(ATR) results in Seckel syndrome', Nature Genetics, 33(4):
497-501.
R5 Iles, N., Rulten, S., El-Khamisy, S.F. and Caldecott, K.W.
(2007) `APLF (C2orf13) is a novel
human protein involved in the cellular response to chromosomal DNA strand
breaks',
Molecular Cellular Biology, 27(10): 3793-803.
R6 Griffith, E., Walker, S., Martin, C., Vagnarelli, P., Stiff,
T., Vernay, B., Al Sanna, N., Saggar, A.,
Hamel, B., Earnshaw, W.C., Jeggo, P.A., Jackson, A.P. and O'Driscoll, M.
(2008) `Mutations
in Pericentrin cause Seckel syndrome with defective ATR-dependent DNA
damage
signalling', Nature Genetics, 40(2): 232-6.
R7 Bicknell, L.S., Walker, S., Klingseisen, A., Stiff, T., Leitch,
A., Kerzendorfer, C., Martin, C.A.,
Yeyati, P., Al Sanna, N., Bober. M., Johnson, D., Wise, C., Jackson, A.P.,
O'Driscoll, M. and
Jeggo, P.A. (2011) `Mutations in ORC1, encoding the largest subunit of the
origin recognition
complex, cause microcephalic primordial dwarfism resembling Meier-Gorlin
syndrome',
Nature Genetics, 43(4): 350-5.
R8 O'Driscoll, M., Cerosaletti, K.M., Girard, P.-M., Dai, Y.,
Stumm, M., Kysela, B., Hirsch, B.,
Gennery, A., Palmer, S.E., Seidel, J., Gatti, R.A., Varon, R., Oettinger,
M.A., Neitzel, H.,
Jeggo, P.A. and Concannon, P. (2001) `DNA Ligase IV mutations identified
in patients
exhibiting development delay and immunodeficiency', Molecular Cell,
8(6): 1175-85.
Outputs can be supplied by the University on request.
Grants supporting the research or awarded based on the research
• Caldecott: Molecular characterisation of single-strand break repair and
related responses and
their role in neuroprotection. MRC Programme, 1 March 2007-28 February
2012; £1,800,000.
• O'Driscoll: How aberrant ataxia telangiestasia and rad3-related (ATR)
pathway function affects
genomic stability. CR-UK Senior Fellowship, 1 July 2007-31 June 2013;
£1,600,000.
• Jeggo: DNA damage responses in mammalian cells and their contribution
to human health
disorders. MRC Programme, 1 October 2006-30 September 2011; £1,498,355.
Details of the impact
As indicated above, the Caldecott/Jeggo/O'Driscoll laboratories have
identified a number of human
genes which, if mutated, result in neurological disease. As a result of
these discoveries, blood and
tissue samples from patients with the relevant disease pathology are
routinely tested for mutations
in these and closely related genes by the Caldecott/Jeggo/O'Driscoll
laboratories, at the request of
clinicians in hospitals within the UK National Health Service (NHS) and
Europe (e.g. Great Ormond
Street, Newcastle General Hospital, Bristol Genetics Laboratory, Nijmegen
Medical Centre), or
within external diagnostic laboratories within the UK and Europe directly
(e.g. Birmingham).
For example, in 2011/2012, based on its identification that APLF and TDP2
are novel human DNA
repair genes, the Caldecott laboratory conducted genetic/biochemical
analyses for defects in
APLF, TDP2, and XRCC1-dependent DNA damage responses in blood samples and
fibroblasts
from patients with neurological/developmental delay, at the request of NHS
clinicians in Bristol (St
Michael's Hospital) [see Section 5, C1] and London (Great Ormond Street)
[C2], and by clinical
geneticists in Nijmegen (Medical Centre). These diagnoses uncovered
mutations in TDP2 as a
cause of epilepsy and spinocerebellar ataxia in four patients [C3].
In like manner, during the current REF period, the Jeggo laboratory has
conducted similar
diagnostic testing at the request of clinicians from a variety of
hospitals in the UK and abroad,
including Great Ormond Street, identifying 11 individuals with RS-SCID out
of 140 who were
screened [C4, C5]. Some of these hospitals have recently established
sequencing procedures/
assays based on guidance from the Jeggo laboratory to aid diagnosis, and
the Jeggo laboratory
continues to advise and carry out a diagnostic service for those patients
in whom sequencing fails
to reveal a mutation or who display a pathology that is suggestive of a
novel or unusual DNA
damage-response-related genetic defect [C5]
In addition, these diagnostic tests are now also conducted externally at
a variety of medical/clinical
genetics centres worldwide, independent of our laboratories, thereby
expanding the reach of our
impact. For example, as a direct result of novel genetic defects
identified by the O'Driscoll/Jeggo
laboratories, multiple clinical genetics centres in the UK (Birmingham,
Great Ormond Street,
Newcastle General Hospital), in North America/Canada (University of
Chicago Genetics Services,
Greenwood Genetics Centre South Carolina, Baylor College of Medicine
Texas, Nemours
Children's Hospital Delaware, and Children's Hospital of Eastern Ontario,
Canada) and in Europe
(Nijmegen Clinical Genetics Center, Dept of Human Genetics Belgium,
Neckar-Enfants Malades
Hospital Paris, Institut für Humangenetik Erlangen and Institut für
Humangenetik Freiburg) provide
molecular diagnostic analyses of causative genes in a clinical genetics
context, and are registered
on the GeneTests service [C6].
As a result of this work, since 2008 the Caldecott/Jeggo/O'Driscoll
laboratories have identified the
underlying genetic defect in more than 150 patients with a range of
hereditary DNA damage-
related disorders. In terms of beneficiaries of this impact, the results
of diagnostic tests are
provided to the relevant clinicians in the form of written reports, which
are employed to inform
additional diagnostic tests and patient care and management. These
diagnoses thus benefit both
the clinical geneticist and the patient; identifying not only the cause of
the patient's disease but also
enabling better disease management. For example, if not first diagnosed,
standard
chemotherapeutic regimes can be fatal in cancer patients who harbour
homozygous TDP2
mutations, and standard conditioning regimes used during bone-marrow
transplantation can be
fatal in LIG4 Syndrome patients. These diagnoses can therefore translate
into increased patient
survival.
Sources to corroborate the impact
Corroborating evidence for Caldecott/O'Driscoll/Jeggo: letters from
clinical geneticists confirming
diagnoses and impact on patient healthcare.
C1 Email from Specialist Registrar (St Michael Hospital).
C2 Email from clinician (Great Ormond Street).
C3 Email from clinical geneticist (Nijmegen Medical Centre).
C4 Emails from Great Ormond Street, Manchester Children's
Hospital, the Beatson Institute and
Gleneagles Medical Centre, Singapore.
C5 Statement from clinician, Peter MacCallum Cancer Centre,
Melbourne, Australia.
C6 http://www.ncbi.nlm.nih.gov/sites/GeneTests/.