TB in Cattle and Badgers: Improving Control of a Multi-species Disease
Submitting Institution
Imperial College LondonUnit of Assessment
Public Health, Health Services and Primary CareSummary Impact Type
PoliticalResearch Subject Area(s)
Medical and Health Sciences: Medical Microbiology, Public Health and Health Services
Summary of the impact
The bovine tuberculosis (TB) research programme led by Professor Donnelly
at Imperial College has been informing policymakers for over a decade.
Professor Donnelly played a leading role in the design, oversight,
analysis and interpretation of the £50 million Randomised Badger Culling
Trial (RBCT, 1998-2006), overseeing a bovine-TB research team at Imperial
since 2001. The RBCT compared two candidate culling policies (large-scale
culling repeated annually and one-off small-scale culls near farms
affected by bovine TB) with areas in which no badger culling took place.
Informed by RBCT results (in which widespread culling decreased cattle
incidence inside the culling area but increased it on neighbouring farms),
the Secretary of State Hilary Benn ruled out badger culling as a control
measure in July 2008. However, the coalition government took a different
view and in December 2011 announced that farmers could apply for licences
to undertake farmer-led (and farmer-funded) badger culling to control TB
in cattle. Several of the government's licensing requirements for badger
culls in England were based on many of the team's results. In contrast,
having proposed in 2010 a large government-led cull, the Welsh government
chose in March 2012 to vaccinate, rather than cull, badgers informed by
the same set of results.
Underpinning research
Key Imperial College London researcher:
Professor Christl Donnelly, Professor of Statistical Epidemiology
(2000-present)
Professor Donnelly played a leading role in the design, oversight,
analysis and interpretation of the Department for Environment, Food and
Rural Affairs (Defra)-funded Randomised Badger Culling Trial (RBCT,
1998-2006) as deputy chair of the Independent Scientific Group on Cattle
TB (ISG). She joined Imperial College in October 2000 and between 2001 and
2007 she led a team of (two to four) government-grant-funded research
associates undertaking RBCT-related research within the Department of
Infectious Disease Epidemiology at Imperial.
The RBCT was very large in scope involving 30 areas of England each of
100sq-km. Each area was randomised to one of three study arms: no culling;
small-scale reactive culling undertaken in response to confirmed infection
found in a cattle herd within the area; and large-scale, annually
repeated, proactive culling which was carried out across the entire
100sq-km area. Badger culls were undertaken within the trial between 1998
and 2005.
The reactive culling strategy was suspended in 2003 when Professor
Donnelly and colleagues found it to be associated with increased TB
incidence in cattle herds (1). Proactive culling continued until 2005 and
in 2006 Professor Donnelly and colleagues found that proactive culling was
associated with significantly decreased TB incidence in cattle herds
within proactively culled trial areas, but significantly increased TB
incidence among cattle herds up to 2km outside proactively culled trial
areas (2, 6).
There were several other strands of complementary research being
undertaken as part of the bovine TB research programme, published in
peer-reviewed scientific journals but also summarised in the Final Report
of the ISG to Ministers (5). Of particular note was the baitmarking study
in which the ranging behaviour of badgers was compared in the three
different types of areas (no culling; reactive culling and proactive
culling). Badger densities were, as expected, highest in the unculled
areas and lowest in the proactive culled areas. Furthermore, badgers in
culled areas (reactive and proactive) and crucially up to 2km outside
proactively culled areas were found to range significantly more widely
than badgers in undisturbed unculled areas (3). Thus, it was hypothesized
that this social "perturbation" to badger populations was responsible for
the increased transmission risks to cattle within reactively culled areas
and up to 2km outside proactively culled areas.
Badgers culled in follow-up proactive culls (i.e. those after the initial
proactive cull) were found to be significantly more likely to be infected
with Mycobacterium bovis, the aetiological agent of bovine TB (4). This is
consistent with the social perturbation caused by recent culling
increasing the risk of badger-to-badger transmission, as well as the risk
of badger-to-cattle transmission.
Professor Donnelly's current TB research (with two post-doctoral
researchers) focuses on modelling and cost-effectiveness analysis of
cattle vaccination in the presence of transmission risks from both badgers
and bought-in cattle. She met with Defra officials in March 2013 to
discuss how the results of current work could inform the government's
planned field trial of a Bacillus Calmette-Guérin (BCG) cattle vaccine.
References to the research
(1) Donnelly, C.A., Woodroffe, R., Cox, D.R., Bourne, J., Gettinby, G.,
Le Fevre, A.M., McInerney, J.P., & Morrison, W.I. (2003). Impact of
localized badger culling on tuberculosis incidence in British cattle. Nature,
426, 834-837. DOI.
Times cited: 136 (as at 30th October 2013 on ISI Web of
Science). Journal Impact Factor: 38.59
(2) Donnelly, C.A., Woodroffe, R., Cox, D.R., Bourne, F.J., Cheeseman,
C.L., Clifton-Hadley, R.S., Gao, Wei, Gettinby, G., Gilks, P., Jenkins,
H., Johnston, W.T., Le Fevre, A.M., McInerney, J.P. & Morrison, W.I.
(2006). Positive and negative effects of widespread badger culling on
tuberculosis in cattle. Nature, 439, 843-846. DOI.
Times cited: 132 (as at 30th October 2013 on ISI Web of
Science). Journal Impact Factor: 38.59
(3) Woodroffe, R., Donnelly, C.A., Cox, D.R., Bourne, F.J., Cheeseman,
C.L., Delahay, R.J., Gettinby, G., McInerney, J.P., & Morrison, W.I.
(2006). Effects of culling on badger Meles meles spatial
organization: implications for the control of bovine tuberculosis. Journal
of Applied Ecology 43, 1-10. DOI.
Times cited: 62 (as at 30th October 2013 on ISI Web of
Science). Journal Impact Factor: 4.74
(4) Woodroffe, R., Donnelly, C.A., Jenkins, H.E., Johnston, W.T., Cox,
D.R., Bourne, F.J., Cheeseman, C.L., Delahay, R.J., Clifton-Hadley, R.S.,
Gettinby, G., Gilks, P., Hewinson, R.G., McInerney, J.P., & Morrison,
W.I. (2006). Culling and cattle controls influence tuberculosis risk for
badgers. Proceedings of the National Academy of Sciences 103,
14713-14717. DOI.
Times cited: 56 (as at 30th October 2013 on ISI Web of
Science). Journal Impact Factor: 9.6
(5) Bourne, J., Donnelly, C., Cox, D., Gettinby, G., McInerney, J.,
Morrison, I., & Woodroffe, R. (2007). Bovine TB: the Scientific
Evidence. A Science Base for a Sustainable Policy to Control TB in Cattle.
Final Report of the Independent Scientific Group on Cattle TB
presented to the Secretary of State for Environment, Food and Rural
Affairs the Rt Hon David Miliband MP. 287pp.
http://webarchive.nationalarchives.gov.uk/20081107201922/http://defra.gov.uk/animalh/tb/isg/pdf/final_report.pdf
(6) Jenkins, H.E., Woodroffe, R., & Donnelly, C.A. (2010). The
Duration of the Effects of Repeated Widespread Badger Culling on Cattle
Tuberculosis Following the Cessation of Culling. PLoS ONE 5(2):
e9090. DOI.
Times cited: 25 (as at 30th October 2013 on ISI Web of
Science). Journal Impact Factor: 3.73
Key funding:
• The Randomised Badger Culling Trial (RBCT) was funded by and
implemented directly by Defra and Defra staff, under advice from the
Independent Scientific Group on Cattle TB (of which Christl Donnelly was
Deputy Chair).
"The RBCT represents nearly 10 years of work (1998-2007) and nearly £50
million of taxpayer investment." according to Defra's Annex B for their
Consultation Document on Badger Culling http://webarchive.nationalarchives.gov.uk/20120616115816/http:/archive.defra.gov.uk/corporate/consult/tb-control-measures/100915-tb-control-measures-annexb.pdf
• Ministry of Agriculture, Fisheries and Food (2002-2004; £140,549),
Principal Investigator (PI) C. Donnelly, Bovine TB in Cattle and Badgers.
• Veterinary Laboratories Agency ( (2002-2004; £124,957), PI C. Donnelly,
Analysis of data derived from Randomised Culling Trial
• Defra (2004-2007; £880,315), PI C. Donnelly, Analysis of data derived
from Randomised Badger Culling Trial.
• Defra (2009-2011; £379,625), PI C. Donnelly, Further analyses of
spatial and temporal trends in the cattle data associated with the
Randomised Badger Culling Trial
• Defra (2011-2014; £481,855), PI C. Donnelly, Transmission modelling and
cost-effectiveness analysis of cattle vaccination at a herd level
Details of the impact
Impacts include: health and welfare; society; public policy and services;
environment; production Main beneficiaries include: UK and Welsh Assembly
Governments, Defra, cattle industry
Bovine TB in cattle remains a high priority for the Defra and one of the
key challenges facing cattle farmers, particularly in the west and south
west of England and in Wales. Although the Krebs Committee (of which
Professor Donnelly was a member) identified in 1997 "compelling" evidence
that badgers were involved in transmitting infection to cattle, it was the
Randomised Badger Culling Trial (RBCT, 1998-2006) which demonstrated
conclusively that badger transmission contributed significantly to the
incidence of bovine TB in cattle and that badger culling could
significantly decrease that incidence. Badger culling is a politically
sensitive issue, and the results of this trial have been critically
examined by various stakeholders. They have undoubtedly stimulated public
debate [1] and have informed Government policy.
In light of the trial results, Defra had considered Government-led (and
Government-funded) culling but their impact assessment showed that the
costs were "too high to justify the benefits achieved in the RBCT" (para
3.26 [2]). Instead the Government proposed that farmers apply for licences
to undertake farmer-led badger culling to control TB in cattle. The
Government [3, 4, 5] used scientific evidence published by Professor
Donnelly and colleagues as the basis for several of their proposed
criteria for culling licenses, including:
- that the culling areas were large (at least 150sq-km, based on
extrapolation of RBCT results);
- that there is land access for culling for >70% of the area (as in
the RBCT);
- that culling in a licensed area would be carried out annually for at
least 4 years (as undertaken in the RBCT proactive areas).
The Government stated that "the RBCT provides the best scientific
evidence available from which to predict the effects of a future culling
policy" [5; see page 1].
The research impacted upon the Welsh government, who similarly proposed
to undertake badger culling in a large area, again based on the RBCT
finding that the overall benefits were increased as the culling area
increased in absolute size ("based on previous experience and studies of
badger culling strategies, including the latest published results of the
Randomised Badger Culling Trial (RBCT), reducing the reservoir of
infection in this way is expected to reduce the number of confirmed cattle
herd breakdowns in the area. There is evidence from the RBCT that 40-50%
of confirmed herd breakdowns in high incidence areas can be attributed to
a local badger source. Although the RBCT differs considerably from the
approach we are proposing for the Intensive Action Area, the analysis of
its results suggests that a sustained reduction in confirmed herd
breakdowns within the area following culling can be achieved.") [6; see
page 5].
Furthermore, the Welsh government proposal followed the RBCT protocol of
culling using cage-trapping and shooting and used RBCT results to predict
the expected minimum impact: "The data from the RBCT shows that there was
a benefit in the cull area but there was a short term increase in bovine
TB in cattle in the surrounding area. Accounting for this, at the end of a
cull and post cull period of 10 years, we expect to have reduced bovine TB
by approximately 22%, overall preventing an estimated 83 confirmed herd
breakdowns that would otherwise have occurred in the absence of culling
badgers in the area." [6; see page 5]
On 20 March 2012 the Welsh Government [7] announced it would launch a
badger vaccination project, rather than a badger culling programme, as
part of its work to eradicate cattle TB from Wales. This decision was
based on the judgement that the effects of culling, informed by RBCT
results, would be not sufficient "to eliminate or substantially reduce the
incidence of bovine TB in cattle" [7].
Sources to corroborate the impact
[1] Examples of public debate:
[2] Defra. The
Government's policy on Bovine TB and badger control in England.
December 2011. (archived
01/11/2013)
[3] Defra. Bovine Tuberculosis: The Government's approach to tackling the
disease and consultation on a badger control policy. September 2010. 54pp
http://webarchive.nationalarchives.gov.uk/20120616115816/http://archive.defra.gov.uk/corporate/consult/tb-control-measures/100915-tb-control-measures-condoc.pdf
(archived
on 01/11/2013) Also of particular interest is Annex B "Scientific Evidence
on Culling"
http://webarchive.nationalarchives.gov.uk/20120616115816/http:/archive.defra.gov.uk/corporate/consult/tb-control-measures/100915-tb-control-measures-annexb.pdf
(archived
on 01/11/2013)
[4] Bovine TB and Badgers — Commons Library Standard Notes. This
information is provided to Members of Parliament in support of their
parliamentary duties. Badgers and Bovine TB: Background. Standard Note
SN/SC/3751 Last updated: 15 February 2011 Author: Dr Elena Ares http://www.parliament.uk/briefing-papers/SN03751
(archived on
01/11/2013) Badger Culling. Standard Note SNSC-5873 Last updated: 16
February 2012 Author: Dr Elena Ares http://www.parliament.uk/briefing-papers/SN05873
(archived on
01/11/2013)
[5] Defra. Bovine TB — Key conclusions from the meeting of scientific
experts, held at Defra on 4th April 2011. From a meeting between Bob
Watson (Chief Scientific Adviser, Defra), Nigel Gibbens (Chief Veterinary
Officer, Defra), Professor Lord John Krebs, Professor Lord Robert May,
Prof Christl Donnelly and 6 other scientists.
http://archive.defra.gov.uk/foodfarm/farmanimal/diseases/atoz/tb/documents/bovinetb-scientificexperts-110404.pdf
(archived
on 01/11/2013)
[6] Welsh Assembly Government. Consultation Document. Bovine TB
Eradication Programme: Consultation on Badger Control in the Intensive
Action Area. 20 September 2010. 14pp. http://wales.gov.uk/docs/drah/consultation/100921badgercontroliaaen.pdf
(archived
on 01/11/2013)
[7] Welsh
Assembly Government. Environment Minister announces programme of
badger vaccination. 20 March 2012. Archived
on 01/11/2013.